We have PDF versions of all of these papers. If you are having trouble finding it on the web, just email us. Undergraduates, grad students, and post-docs affiliated with the lab are denoted with U, G, and P, respectively.

Brown, N.E.M.G, Milazzo, M., Rastrick, S.P.S., Hall-Spencer, J.M., Therriault, T.W. & Harley, C.D.G. 2018. Natural acidification changes the timing and rate of succession, alters community structure, and increases homogeneity in marine biofouling communities. Global Change Biology 24:e112-e127.

KordasG, R.L., Donahue, I., Harley, C.D.G. 2017. Herbivory enables marine community to resist warming. Science Advances. 3:e1701349.

Harley, C.D.G., Connell, S. D., Doubleday, Z. A., Kelaher, B., Russell, B.D., Sara, G., Helmuth, B. 2017. Conceptualizing ecosystem tipping points within a physiological framework. Ecology and Evolution 7:6035-6045.

Proum, S., Harley, C.D.G., Steele, M., & Marshall, D.J. 2017. Aerobic and behavioural flexibility allow estuarine gastropods to flourish in rapidly changing and extreme pH conditions. Marine Biology 5:97.

Kroeker, K.J., KordasG, R.L., & Harley, C.D.G. 2017. Embracing interactions in ocean acidification research: confronting multiple stressor scenarios and context dependence. Biology Letters 13:20160802.

Connell, S.D., Doubleday, Z.A., Hamlyn, S.B., Foster, N.R., Harley, C.D.G., Helmuth, B., Kelaher, B.P., Nagelkerken, I., Sara, G., & Russell, B.D. 2017. How ocean acidification can benefit calcifiers. Current Biology 27:R95-R96.

Harley, C.D.G. 2016 Phycology for the ecologist. Journal of Phycology 52:898-900.

Sunday, J.M., Fabricius, K.E., Kroeker, K.J., AndersonG, K.M., BrownG, N.E., Barry, J.P., Connell, S.D., Dupont, S., Gaylord, B., Hall-Spencer, J.M., Klinger, T., Milazzo, M., Munday, P.L., Russell, B.D., Sanford, E., Thiyagarajan, V., VaughanG. M.L.H., Widdicombe, S., & Harley, C.D.G. 2016. Ocean acidfication can mediate biodiversity shifts by changing biogenic habitat. Nature Climate Change 7:81-85.

Sinclair, B.J., MarshallP, K.E., Sewell, M.A., Levesque, D.L., Willett, C.S., Slotsbo, S., Dong, Y., Harley, C.D.G., Marshall, D.J., Helmuth, B.S., & Huey, R.B. 2016. Can we predict ectotherm responses to cliamte change using thermal performance curves and body temperatures? Ecology Letters 19:1372-1385.

BrownG, N.E.M., Therriault, T.W., & Harley, C.D.G. 2016. Field-based experimental acidification alters fouling community structure and reduces diversity. Journal of Animal Ecology 85:1328-1339.

KordasG, R.L., & Harley, C.D.G. 2016. Demographic responses of coexisting species to in situ warming. Marine Ecology Progress Series. 546:147-161.

Helmuth, B., Choi, F., Matzelle, A., Torossian, J.L., Morello, S.L., Mislan, K.A.S., Lauren, Y., Strickland, D., Szathmary, P.L., Gilman, S.E., Tockstein, A., Hilbish, T.J., Burrows, M.T., Power, A.M., Gosling, E., Mieszkowska, N., Harley, C.D.G., Nishizaki, M., Carrington, E., Menge, B., Petes, L., Foley. M.M., Johnson, A., Poole, M., Noble, M.M., Richmond, E.L., Robart, M., Robinson, J.S., Sones, J., Broitman, B.R., Denny, M.W., Mach, K.J., Miller, L.P., O’Donnell, M., Ross, P., Hofmann, G.E., Zippay, M., Blanchette, C., Macfarlan, J.A., Carpizo-Ituarte, E., Ruttenberg, B., Mejia, C.E.P., McQuaid, C.D., Lathlean, J., Monaco, C.J., Nicastro, K.R., & Zardi, G. 2016. Long-term, high frequency in situ measurements of intertidal mussel bed temperatures using biomimetic sensors. Scientific Data 3:160087.

Krumhansl, K. A., Demes,G K.W., Carrington, E., & Harley, C.D.G. 2015. Divergent growth strategies between red algae and kelps influence biomechanical properties. American Journal of Botany 102:1938-1944.

KordasG, R.L., Dudgeon, S., Storey, S., & Harley, C.D.G. 2015. Intertidal community responses to field-based experimental warming. Oikos 124:888-898.

GoodingG, R.A. & Harley, C.D.G. 2015. Quantifying the effects of predator and prey body size on sea star feeding behaviours. Biological Bulletin 228:192-200.

DeLong, J.P., Gilbert, B., Shurin, J.B., Savage, V.M., Barton, B.T., Clements, C.F., Dell, A.I., Greig, H.S., Harley, C.D.G., Kratina, P., McCann, K.S., Tunney, T.D., Vasseur, D.A., & O’Connor, M.I. 2015. The body size dependence of trophic cascades. American Naturalist 185:354-366.

Gaylord, B., Kroeker, K.J., Sunday, J.M., AndersonG, K.M., Barry, J.P., BrownG, N.E., Connell, S.D., Dupont, S., Fabricius, K.E., Hall-Spencer, J.M., Klinger, T., Milazzo, M., Munday, P.L., Russell, B.D., Sanford, E., Schreiber, S.J., Thiyagrajan, V., VaughanG, M.L.H., Widdicombe, S., & Harley, C.D.G. 2015. Ocean acidification through the lens of ecological theory. Ecology 96:3-15.

Helmuth, B., Russell, B.D., Connell, S.D., Dong, Y., Harley, C.D.G., Lima, F.P. , Sara, G., Williams, G.A., & Mieszkowska, N. 2014. Beyond long-term averages: making biological sense of a rapidly changing world. Climate Change Responses 1:6.

Gilbert, B., Tunney, T.D., McCann, K.S., DeLong, J.P., Vasseur, D.A., Savage, V., Shurin, J.B., Dell, A.I., Barton, B.T., Harley, C.D.G., Kharouba, H.M., Kratina, P., Blanchard, J.L., Clements, C., Winder, M., Greig, H.S., & O’Connor, M.I. 2014. A bioenergetics framework for the temperature dependence of trophic interaction strength. Ecology Letters 17:902-914.

Vasseur, D.A., DeLong, J.P., Gilbert, B., Grieg, H., Harley, C.D.G., McCann, K.S., O’Connor, M.I., Tunney, T., & Savage, V. 2014. Temperature variation puts species at greater risk under global change. Proceedings of the Royal Society B 281:20132612.

DemesG, K.W., Harley, C.D.G., Anderson, L.M., & Carrington, E. 2013. Shifts in morphological and mechanical traits compensate for performance costs of reproduction in a wave-swept seaweed. Journal of Ecology 101:963-970.

DemesG, K.W., Pruitt, J.N., Harley, C.D.G., & Carrington, E. 2013. Survival of the weakest: Increased frond mechanical strength in a wave-swept kelp inhibits self-pruning and increases mortality. Functional Ecology 27:439-445. [Winner of the 2013 Haldane Prize for best paper by a young author published in Functional Ecology.]

Harley, C.D.G., AndersonG, K.M., LebretonU, C.A.-M., MacKayU, A., Ayala-Díaz, M., ChongU, S.L., PondU, L.M., Amerongen MaddisonU, J.H., HungU, B.H.C., IversonU, S.L., & WongU, D.C.M. 2013. The introduction of Littorina littorea to British Columbia, Canada: potential impacts and the importance of biotic resistance. Marine Biology 160:1529-1541.

SinghG, G., Markel, R., MartoneP, R., Salomon, A., Harley, C.D.G., & Chan, K. 2013. Sea otters homogenize mussel beds and reduce habitat provisioning in a rocky intertidal ecosystem. PLoSONE 8(5):e65435.

DemesG, K., KordasG, R., & JorveG, J. 2012. Ferry wakes increase seaweed richness and abundance in a sheltered rocky intertidal habitat. Hydrobiologia 693:1-11.

Harley, C.D.G., AndersonG, K.M., DemesG, K.W., JorveG, J.P., KordasG, R.L., CoyleU, T.A., & Graham, M.H. 2012. Effects of climate change on global seaweed communities. Journal of Phycology 48:1064-1078.

Russell, B.D., Harley, C.D.G., Wernberg, T., Mieszkowska, N., Widdicombe, S., Hall-Spencer, J.M., & Connell, S.D. 2012. Predicting ecosystem shifts requires new approaches that integrate the effects of climate change across entire systems. Biology Letters 8:164-166.

CrimG, R.N., Sunday, J.M., & Harley, C.D.G. 2011. Elevated seawater CO2 concentrations impair larval development and reduce larval survival in endangered northern abalone (Haliotis kamtschatkana). Journal of Experimental Marine Biology and Ecology 400:272-277.

Harley, C.D.G. 2011. Climate change, keystone predation, and biodiversity loss. Science 334:1124-1127.

Harley, C.D.G. & O’RileyU 2011. Non-linear density dependent effects of an intertidal ecosystem engineer. Oecologia 166:531-541.

KordasG, R.L., Harley, C.D.G., & O’Connor, M.I. 2011. Community ecology in a warming world: the influence of temperature on interspecific interactions. Journal of Experimental Marine Biology and Ecology 400:218-226.

Reuter, K.E., Lotterhos, K.E., CrimG, R.N., Thompson, C.A., & Harley, C.D.G. 2011. Ocean acidification increases sperm limitation and risk of polyspermy in the red sea urchin Strongylocentrotus franciscanus. Global Change Biology 17:163:171.

Sunday, J.M., CrimG, R., Harley, C.D.G., & Hart, M.W. 2011. Quantifying rates of evolutionary adaptation in response to ocean acidification. PLoS One 6(8):e22881.

Kroeker, K.J., KordasG, R.L., CrimG, R.N., & SinghG, G.G. 2010. Meta-analysis reveals negative yet variable effects of ocean acidification on marine organisms. Ecology Letters 13:1419-1434.

BatesP, A., Stickle, W., & Harley, C.D.G. 2010. Impact of temperature on the parasitic association between a sperm-feeding scuticociliate and Pacific west coast sea stars. Journal of Experimental Marine Biology and Ecology 384:44-50.

NienhuisG, S., Palmer, A.R., & Harley, C.D.G. 2010. Elevated CO2 affects shell dissolution rate but not calcification rate in a marine snail. Proceedings of the Royal Society B 277:2553-2558.

BatesP, A., Hilton, B.J., & Harley, C.D.G. 2009. Effects of seasonal variation and temperature on wasting disease in the keystone predatory sea star, Pisaster ochraceus. Diseases of Aquatic Organisms 86:245-251.

Denny, M.W., Hunt, L.J.H., Miller, L.P., & Harley, C.D.G. 2009. On the prediction of extreme ecological events. Ecological Monographs 79:397-421.

GoodingG, B., Harley, C.D.G., & TangU, E. 2009. Multiple climate variables increase growth and feeding rates of a keystone predator. Proceedings of the National Academy of Sciences USA 106:9316-9321.

Harley, C.D.G., Denny, M.W., Mach, K., & Miller, L.P. 2009. Thermal stress and morphological adaptations in limpets. Functional Ecology 23:292-301.

Harley, C.D.G., & Paine, R.T. 2009. Contingencies and compounded rare perturbations dictate sudden distributional shifts during periods of gradual climate change. Proceedings of the National Academy of Sciences USA 106:11172-11176.

HeldU, M., & Harley, C.D.G. 2009. Responses to low salinity by the sea star Pisaster ochraceus from high- and low-salinity populations. Invertebrate Biology 128:381-390.

Miller, L.P., Harley, C.D.G., & Denny, M.W. 2009. The role of temperature and desiccation stress in limiting the local-scale distribution of the owl limpet, Lottia gigantea. Functional Ecology 23:756-767.

Harley, C.D.G. 2008. Tidal dynamics, topography, and temperature-mediated mass mortalities on rocky shores. Marine Ecology Progress Series 371:37-46.

MorelissenG, B., & Harley, C.D.G. 2007. The effects of thermal stress on producers, consumers, and plant-herbivore interactions in an intertidal community. Journal of Experimental Marine Biology and Ecology 348:162-173.

Denny, M.W., & Harley, C.D.G. 2006. Hot limpets: predicting body temperature in a conductance-mediated thermal system. Journal of Experimental Biology 209:2409-2419

Denny, M.W., Miller, L.P., & Harley, C.D.G. 2006. Thermal stress on intertidal limpets: long-term hindcasts and lethal limits. Journal of Experimental Biology 209:2420-2431.

Gilman, S.E., Harley, C.D.G., Strickland, D.C., Vanderstraeten, O., O’Donnell, M.J., & Helmuth, B. 2006. Evaluation of effective shore level as a method of characterizing intertidal wave exposure regimes. Limnology & Oceanography Methods 4:448-457.

Harley, C.D.G. 2006. Direct and indirect effects of ecosystem engineering and herbivory on intertidal community structure. Marine Ecology Progress Series 317:29-39.

Harley, C.D.G., Hughes, A.R., Hultgren, K.M., Miner, B.G., Sorte, C.J.B., Thornber, C.S., Rodriguez, L.F., Tomanek, L., & Williams, S. L. 2006. The impact of climate change in coastal marine systems. Ecology Letters 9:228-241.

Harley, C.D.G., Pankey, M.S., Wares, J.P., Wonham, M.J., & Grosberg, R. 2006. Color polymorphism and genetic structure in the sea star Pisaster ochraceus. Biological Bulletin 211:248-262.

Helmuth, B., Broitman, B., Blanchette, C.A., Gilman, S., Halpin, P., Harley, C.D.G., O’Donnell, M., Hofmann, G.E., Menge, B., & Strickland, D. 2006. Mosaic patterns of thermal stress in the rocky intertidal zone: implications for climate change. Ecological Monographs 76:461-479.

Wonham, M.J., O’ConnorU, M., & Harley, C.D.G. 2005. Enhancement of introduced and native species by a dominant invader. Marine Ecology Progress Series 289:109-116.

Harley, C.D.G. & Rogers-Bennett, L. 2004. The potential synergistic effects of climate change and fishing pressure on exploited invertebrates on rocky intertidal shores. California Cooperative Oceanic Fisheries Investigations Reports 45:98-110.

Denny, M.W., Helmuth, B., Leonard, G.H., Harley, C.D.G., Hunt, L.J.H., & Nelson, E.K. 2004. Quantifying scale in ecology: lessons from a wave-swept shore. Ecological Monographs 74:513-532.

Harley, C.D.G. 2003. Abiotic stress and herbivory interact to set range limits across a two-dimensional stress gradient. Ecology 84:1477-1488.

Harley, C.D.G. & Helmuth, B.S.T. 2003. Local- and regional-scale effects of wave exposure, thermal stress, and absolute vs. effective shore level on patterns of intertidal zonation. Limnology & Oceanography 48:1498-1508.

Harley, C.D.G. & LopezU, J.P. 2003. The natural history, physiology, and ecological impacts of the intertidal mesopredators, Oedoparena spp. (Diptera, Dryomyzidae). Invertebrate Biology 122:61-73. 

Harley, C.D.G., Smith, K.F., & MooreU, V.L. 2003. Environmental variability and biogeography: the relationship between bathymetric distribution and latitudinal range size in marine algae and gastropods. Global Ecology and Biogeography 12:499-506.