Final exams may be viewed on Jan. 12 (2pm) or Jan.16 (1:30pm) in my office.

Grad presentations: Kenny Askelson on detecting local adaptation; Sierra Sullivan on introgression; Yee Fay Lim on time-dependent molecular rates; Maddie Ore on supergenes; Freek de Haas on Muller's ratchet; Casandra Elphinstone on rates of evolution; Ryan Stratychuk on pop. gen. of clonality; Seth Watt on population genetics of sex; Takuji Usui on evolution of range limits; Tal Shalev on complex traits;

The exam for this calss has been scheduled for December 5 at 3:30 pm in WOODWARD 5.

Midterm key

Most of my lecture notes are available as a pdf. Warning: we will add topics and vary the order from these notes fairly often.


Assignment 7: Due in class Thursday Nov. 23:Note two-week gapAnswers

Assignment 6: Due in class Thursday Nov. 9:Answers

Assignment 5: SKIP QUESTION 2! Due in class Thursday Oct.19: Answers

Assignment 4: Due in class Thursday Oct.12: Answers

Assignment 3: Due in class Thursday Oct.5: : Answers

Assignment 2: Due in class Thursday September 28: Answers

Assignment 1: Due in class Thursday September 21: Answers

Helpful materials

Approximate course outline

Formula sheet

Old midterm; another previous midterm, Midterm 2008 answer key, Midterm 2009, Midterm 2010, midterm 2014, and and an old final, with its answers .


Prof. Michael Whitlock

Office: Biodiversity 216

Office Hours: Mondays 1:30-2:30 (and after class)



Classes are in room 405 of the West Mall Swing Space, Tuesdays and Thursdays 11:00-12:20.

The midterm will be on either October 19 or October 26, determined by majority vote of the class.

Assignments are due in class.

The (optional) textbook is Charlesworth and Charlesworth, The Elements of Population Genetics. You may prefer an equally good (in different ways) text by Joe Felsenstein, which is freely available as a PDF at

There are also many references on reserve in Woodward Library, including copies of Hartl and Clark.

The final grade for BIOL 434 will be composed of 50% final exam, 30% midterm, and 20% assignment marks and class participation.

Make-up policy: Make-ups for the final exam will follow the university policy. No make-ups for the mid-term are given. If a student misses the mid-term for a valid reason, the final exam grade will be counted for the mid-term as well.

Some quantitative genetic practice questions are here. (answers)