Throughout the term, any important announcements will be listed here.

The answer key for the midterm is here.

Most of my lecture notes are available as a pdf. Warning: we will add topics and vary the order from these notes fairly often.


No assignment due next week. We'll have a double assignment due the next week.

Assignment 6: answers

Assignment 5: answers

Assignment 4: answers

Assignment 3: answers

Assignment 2: answers

Assignment 1: answers

Helpful materials

Approximate course outline

Formula sheet

Old midterm; another previous midterm, Midterm 2008 answer key, Midterm 2009, Midterm 2010, midterm 2014, and and an old final, with its answers . Midterm key from last year


Prof. Michael Whitlock

Office: Biodiversity 216

Office Hours: Mondays 1:30-2:30 (and after class)



Classes are in room 409 of the West Mall Swing Space, Tuesdays and Thursdays 11:00-12:20.

The midterm will be on October 25, determined by majority vote of the class.

Assignments are due in class.

The (optional) textbook is Charlesworth and Charlesworth, The Elements of Population Genetics. You may prefer an equally good (in different ways) text by Joe Felsenstein, which is freely available as a PDF at

There are also many references on reserve in Woodward Library, including copies of Hartl and Clark.

The final grade for BIOL 434 will be composed of 50% final exam, 30% midterm, and 20% assignment marks and class participation.

Make-up policy: Make-ups for the final exam will follow the university policy. No make-ups for the mid-term are given. If a student misses the mid-term for a valid reason, the final exam grade will be counted for the mid-term as well.

Some quantitative genetic practice questions are here. (answers)