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  1. Harter T. S., L. T. N. Heinsbroek, J. W. Schrama 2014. The source of dietary non-protein energy affects in vivo protein digestion in African catfish (Clarias gariepinus). In Press
  2. Harter T. S., R. B. Shartau, D. W. Baker, D. C. Jackson, A. L. Val and C. J. Brauner 2014. Preferential intracellular pH regulation represents a general pattern of pH homeostasis during acid-base disturbances in the armoured catfish, Pterygoplichthys pardalis. DOI 10.1007/s00360-014-0838-8
  3. Harter T., R. B. Shartau, C. J. Brauner, A. P. Farrell 2014. Portable, fast and inaccurate: validation of the i-STAT system for the analysis of blood parameters in fish. In Press
  4. Harter, T. S., J. A. J. Verreth, L. T. H. Heinsbroek and J. W. Schrama 2013. Isoenergetic replacement of fat by starch in diets for African catfish (Clarias gariepinus): Effect on water fluxes in the gastro intestinal tract. PLoS ONE 8 (1):e55245
  5. Harter, T., P. Bossier, J. Verreth, S. Bodé, D. Ha, A.-E. Debeer, N. Boon, P. Boeckx, W. Vyverman, N. Nevejan 2012. Carbon and nitrogen mass balance during flue gas treatment with Dunaliella salina cultures. Journal of Applied Phycology 25 (2):359-368


Till Harter

PhD Student

Web page: Lab page
Research area: Comparative Physiology
Supervisor: C. Brauner
History: BSc. Agricultural Sciences at Hohenheim University, Germany
MSc. Aquaculture and Fisheries at Wageningen University, The Netherlands

Osmoregulation in marine fishes largely takes place in the intestine and is a highly energy and oxygen demanding process. However, the blood supply to this organ is variable and may be largely reduced during exercise or stress. At the same time these situations are known to impose an osmotic disturbance to the animal and the oxygen demand for osmoregulation at the intestine will increase. Some teleost fishes may have evolved a system to actively unload oxygen from Root-effect hemoglobins and thereby safeguard oxygen supply to the intestine (and potentially other vital tissues) and maintain osmotic balance during stress. A key aspect of this system is the presence of plasma accessible carbonic anhydrase in the vasculature of the intestine and my goal is to provide functional in vitro and in vivo evidence for the presence and function of this enzyme.



Four Year Fellowships (FYF) For PhD Students

For Research


International Tuition Award

For Research

Last updated 28 April 2014